Tolypelepidida is an extinct order of heterostracan vertebrates. These armored jawless fish superficially resemble their relatives, the cyathaspids, though, researchers place the tolypelepids as a sister group to the cyathaspids and the pteraspidids (and both groups' daughter taxa). A recent study by Lundgren and Blom in 2013 implies that the order is paraphyletic, with the type genus, Tolypelepis, being the sister taxon of Cyathaspidiformes.
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