Gustav Georg Embden (November 10, 1874 – July 25, 1933) was a German chemist.He taught at the University of Frankfurt am Main from its founding in 1914. He conducted studies on carbohydrate metabolism and muscle contraction, and was the first to discover and link together all the steps involved in the conversion of glycogen to lactic acid. In 1918 Otto Fritz Meyerhof explained – cellular metabolism by showing that it involved the breakdown of glucose to lactic acid.